The characters from the animated Madagascar franchise, Alex, Melman, Marty and Gloria celebrate love in this Valentine special.

The Madagascar animals fly back to New York City, but crash-land on an African nature reserve, where they meet others of their own kind, and Alex especially discovers his royal heritage as prince of a lion pride.

Robin Monroe, a New York magazine editor, and the gruff pilot Quinn Harris must put aside their mutual dislike if they are to survive after crash landing on a deserted South Seas island.